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FMA treats them as spaces.
In NIF GA they are perhaps implicitly structures (superficial feature implies that it is something physical not abstract)
If we ever want to talk about genes expressed in a sulcus they cannot be immaterial.
But this would probably be unusual.
One possibility is to have parallel hierarchies for the feature and the space (this would not quite parallel sulcus vs gyrus). Best avoided.
I am tending towards the FMA classification.
Required for issue #314
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